Research Funding

  • August 1, 2004 - December 31, 2025 - David W. Smith Workshop on Malformations and Morphogenesis , Co-Principal Investigator . Sponsor: NIH, Sponsor Award ID: R13HD048170
  • June 1, 2019 - May 31, 2024 - Precision models of ARX-associated neurodevelopmental disorders , Co-Principal Investigator . Sponsor: NIH, Sponsor Award ID: R01NS113516
  • May 1, 2018 - April 30, 2022 - Arx Associated Transcriptional Networks in Neocortical Development , Principal Investigator . Sponsor: NIH, Sponsor Award ID: R01NS100007
  • May 1, 2017 - April 30, 2018 - Arx Associated Transcriptional Networks in Neocortical Development , Principal Investigator . Sponsor: NIH, Sponsor Award ID: R56NS100007
  • July 1, 2003 - April 30, 2015 - The Role of ARX in Normal and Abnormal Brain Development , Principal Investigator . Sponsor: NIH, Sponsor Award ID: R01NS046616
  • February 15, 2003 - August 31, 2012 - Non-Radial Cell Migration in CNS Development , Principal Investigator . Sponsor: NIH, Sponsor Award ID: R56NS045034
  • February 15, 2003 - August 31, 2011 - Non-Radial Cell Migration in CNS Development , Principal Investigator . Sponsor: NIH, Sponsor Award ID: R01NS045034
  • May 15, 2000 - April 30, 2003 - CELL MIGRATION AND CNS DEVELOPMENT ANOMALIES , Principal Investigator . Sponsor: NIH, Sponsor Award ID: R21NS039949
  • July 1, 1993 - June 30, 1998 - LINEAGE ANALYSIS OF THE THALAMUS , Principal Investigator . Sponsor: NIH, Sponsor Award ID: K08NS001664

Education

Residency, - School of Medicine, University of California, San Francisco

Selected Publications

  1. Graunke H, Saunders R, McBride T, Golden J, Kuipers EJ, McClellan M, Zimlichman E, Kahn CH. Strategies to Ensure a Resilient Health Care Workforce: International Models on Health Professions. NEJM Catal Innov Care Deliv. 2026 Apr; 7(4):CAT250263. View on PubMed
  2. Mohanty RP, Calabrese DR, Moghbeli K, Iasella CJ, Devonshire K, Zeng A, Chen K, Singer JP, Hays SR, Leard LE, Estrada AV, Shah RJ, Kolaitis NA, Golden JA, Langelier CR, McDyer JF, Greenland JR. Airway molecular signatures in antibody-mediated lung transplant rejection. Am J Transplant. 2026 Feb 12. View on PubMed
  3. Lim Y, Akula SK, Myers AK, Chen C, Rafael KA, Ibach MG, Trevathan E, Walsh CA, Golden JA, Cho G. ARX mutation-associated interneuron defects provide insights into mechanisms underlying developmental epilepsies. Brain. 2026 Feb 03. View on PubMed
  4. Myers AK, Sakheim M, Rivell C, Fengler C, Festa LK, Guerra KM, Jarrahy L, Shin R, Case M, Chapman C, Basel L, Springer S, Kern N, Gidicsin J, Cho G, Kim S, Tighiouart M, Golden JA. Anxiety-Associated Behaviors Following Ablation of Miro1 from Cortical Excitatory Neurons. eNeuro. 2026 01; 13(1). View on PubMed
  5. Lin SY, Tsai PC, Su FY, Chen CY, Li F, Zhao J, Ho YY, Lee TM, Healey E, Lin PJ, Kao TW, Vremenko D, Roetzer-Pejrimovsky T, Sholl L, Dillon D, Lin NU, Meredith D, Ligon KL, Lo YC, Chaisuriya N, Cook DJ, Woehrer A, Meyerhardt J, Ogino S, Nasrallah MP, Golden JA, Signoretti S, Chiang JH, Yu KH. Contrastive learning enhances fairness in pathology artificial intelligence systems. Cell Rep Med. 2025 Dec 16; 6(12):102527. View on PubMed
  6. Lim Y, Norman M, Chen C, Cho IT, Walt DR, Cho G, Golden JA. Single-Molecule Array (Simoa) Technology as a Highly Sensitive Screening Tool for Protein-Protein Interactions. ACS Sens. 2025 Nov 28; 10(11):8256-8261. View on PubMed
  7. Zhao J, Lin SY, Attias R, Mathews L, Engel C, Larghero G, Vremenko D, Kao TW, Lee TH, Wang YH, Tsai CC, Marostica E, Lo YC, Meredith D, Ligon KL, Arnaout O, Roetzer-Pejrimovsky T, Lin SC, Shih NN, Chaisuriya N, Cook DJ, Chiang JH, Liu CJ, Woehrer A, Golden JA, Nasrallah MP, Yu KH. Uncertainty-aware ensemble of foundation models differentiates glioblastoma from its mimics. Nat Commun. 2025 Sep 29; 16(1):8341. View on PubMed
  8. Wang X, Zhao J, Marostica E, Yuan W, Jin J, Zhang J, Li R, Tang H, Wang K, Li Y, Wang F, Peng Y, Zhu J, Zhang J, Jackson CR, Zhang J, Dillon D, Lin NU, Sholl L, Denize T, Meredith D, Ligon KL, Signoretti S, Ogino S, Golden JA, Nasrallah MP, Han X, Yang S, Yu KH. A pathology foundation model for cancer diagnosis and prognosis prediction. Nature. 2024 10; 634(8035):970-978. View on PubMed
  9. Cohen ML, Brumwell AN, Ho TC, Garakani K, Montas G, Leong D, Ding VW, Golden JA, Trinh BN, Jablons DM, Matthay MA, Jones KD, Wolters PJ, Wei Y, Chapman HA, Le Saux CJ. A fibroblast-dependent TGF-β1/sFRP2 noncanonical Wnt signaling axis promotes epithelial metaplasia in idiopathic pulmonary fibrosis. J Clin Invest. 2024 Jul 09; 134(18). View on PubMed
  10. Lim Y, Akula SK, Myers AK, Chen C, Rafael KA, Walsh CA, Golden JA, Cho G. ARX regulates cortical interneuron differentiation and migration. bioRxiv. 2024 Jun 06. View on PubMed
  11. Aguilar OA, Qualls AE, Gonzalez-Hinojosa MDR, Obeidalla S, Kerchberger VE, Tsao T, Singer JP, Looney MR, Raymond W, Hays SR, Golden JA, Kukreja J, Shaver CM, Ware LB, Christie J, Diamond JM, Lanier LL, Greenland JR, Calabrese DR. MICB Genomic Variant Is Associated with NKG2D-mediated Acute Lung Injury and Death. Am J Respir Crit Care Med. 2024 01 01; 209(1):70-82. View on PubMed
  12. Santos J, Wang P, Shemesh A, Liu F, Tsao T, Aguilar OA, Cleary SJ, Singer JP, Gao Y, Hays SR, Golden JA, Leard L, Kleinhenz ME, Kolaitis NA, Shah R, Venado A, Kukreja J, Weigt SS, Belperio JA, Lanier LL, Looney MR, Greenland JR, Calabrese DR. CCR5 drives NK cell-associated airway damage in pulmonary ischemia-reperfusion injury. JCI Insight. 2023 11 08; 8(21). View on PubMed
  13. Greenland JR, Guo R, Lee S, Tran L, Kapse B, Kukreja J, Hays SR, Golden JA, Calabrese DR, Singer JP, Wolters PJ. Short airway telomeres are associated with primary graft dysfunction and chronic lung allograft dysfunction. J Heart Lung Transplant. 2023 12; 42(12):1700-1709. View on PubMed
  14. Santos J, Hays SR, Golden JA, Calabrese DR, Kolaitis N, Kleinhenz ME, Shah R, Estrada AV, Leard LE, Kukreja J, Singer JP, Greenland JR. Decreased Lymphocytic Bronchitis Severity in the Era of Azithromycin Prophylaxis. Transplant Direct. 2023 Sep; 9(9):e1495. View on PubMed
  15. Cohen ML, Brumwell AN, Che Ho T, Montas G, Golden JA, Jones KD, Wolters PJ, Wei Y, Chapman HA, Le Saux CJ. A fibroblast-dependent TGFβ1/sFRP2 noncanonical Wnt signaling axis underlies epithelial metaplasia in idiopathic pulmonary fibrosis. bioRxiv. 2023 Aug 04. View on PubMed
  16. Nguyen A, Chen J, Isaza E, Panchal N, Deiter F, Hoover J, Trinh B, Hays SR, Golden JA, Singer JP, Brzezinski M, Greenland JR, Kukreja J. Biofire pneumonia panel in lung donors: faster detection but limited pathogens. Transpl Infect Dis. 2023 Aug; 25(4):e14091. View on PubMed
  17. Nasrallah MP, Zhao J, Tsai CC, Meredith D, Marostica E, Ligon KL, Golden JA, Yu KH. Machine learning for cryosection pathology predicts the 2021 WHO classification of glioma. Med. 2023 08 11; 4(8):526-540.e4. View on PubMed
  18. Singer JP, Christie JD, Diamond JM, Anderson MA, Benvenuto LA, Gao Y, Arcasoy SM, Lederer DJ, Calabrese D, Wang P, Hays SR, Kukreja J, Venado A, Kolaitis NA, Leard LE, Shah RJ, Kleinhenz ME, Golden J, Betancourt L, Oyster M, Zaleski D, Adler J, Kalman L, Balar P, Patel S, Medikonda N, Koons B, Tevald M, Covinsky KE, Greenland JR, Katz PK. Development of the Lung Transplant Frailty Scale (LT-FS). J Heart Lung Transplant. 2023 07; 42(7):892-904. View on PubMed
  19. Singer JP, Calfee CS, Delucchi K, Diamond JM, Anderson MA, Benvenuto LA, Gao Y, Wang P, Arcasoy SM, Lederer DJ, Hays SR, Kukreja J, Venado A, Kolaitis NA, Leard LE, Shah RJ, Kleinhenz ME, Golden J, Betancourt L, Oyster M, Brown M, Zaleski D, Medikonda N, Kalman L, Balar P, Patel S, Calabrese DR, Greenland JR, Christie JD. Subphenotypes of frailty in lung transplant candidates. Am J Transplant. 2023 04; 23(4):531-539. View on PubMed
  20. Odisho AY, Liu AW, Maiorano AR, Bigazzi MOA, Medina E, Leard LE, Shah R, Venado A, Perez A, Golden J, Kleinhenz ME, Kolaitis NA, Maheshwari J, Trinh BN, Kukreja J, Greenland J, Calabrese D, Neinstein AB, Singer JP, Hays SR. Design and implementation of a digital health home spirometry intervention for remote monitoring of lung transplant function. J Heart Lung Transplant. 2023 06; 42(6):828-837. View on PubMed

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